BSc, The Hebrew University of Jerusalem, Biology, 1985
MSc, The Hebrew University of Jerusalem, Human Genetics,1988
PhD, The Hebrew University of Jerusalem, Genetics,1994
PostDoc, The Hebrew University of Jerusalem, Genetics,1995-1998


An Sci 875 Molecular Aspects of Animal Breeding
AnSci/DySci 361 Intro to Animal & Veterinary Genetics
AnSci/DySci 362 Veterinary Genetics
AnSci/DySci 363 Principles of Animal Breeding


Integrative analysis of methylomic and transcriptomic data in fetal sheep muscle tissues in response to maternal diet during pregnancy. Namous H, Peñagaricano F, Del Corvo M, Capra E, Thomas DL, Stella A, Williams JL, Marsan PA,Khatib H BMC Genomics 2018, 19:123 | Published on: 6 February 2018

Gross N, Kropp J, Khatib H. 2017. MicroRNA Signaling in Embryo Development. Biology (Basel)6(3). pii: E34. Review.

Cole JB, Bormann JM, Gill CA, Khatib H, Koltes JE, Maltecca C, Miglior F. 2017. BREEDING AND GENETICS SYMPOSIUM: Resilience of livestock to changing environments. J Anim Sci. 95(4):1777-1779.

Gross N, Kropp J, and Khatib H . Sexual Dimorphism of miRNAs Secreted by Bovine In vitro-produced Embryos. (2017) Front. Genet. 8:39

Kropp J, Carrillo JA, Namous H, Daniels A, Salih SM, Song J, and Khatib H. Male fertility status is associated with DNA methylation signatures in sperm and transcriptomic profiles of bovine preimplantation embryos. (2017)BMC Genomics 18:280

Kropp J, Khatib H. 2015. mRNA fragments in in vitro culture media are associated with bovine preimplantation embryonic development. Front Genet. 2015 Aug 24;6:273.

Peñagaricano F, Valente BD, Steibel JP, Bates RO, Ernst CW, Khatib H, Rosa GJ. 2015. Exploring causal networks underlying fat deposition and muscularity in pigs through the integration of phenotypic, genotypic and transcriptomic data. BMC Syst Biol. 2015 Sep 16;9:58.

The FAANG Consortium, L Andersson, AL Archibald, CD Bottema, R Brauning, SC Burgess, DW Burt, E Casas, HH Cheng, L Clarke, C Couldrey, BP Dalrymple, CG Elsik, S Foissac, E Giuffra, MA Groenen, BJ Hayes, LS Huang, H Khatib, JW Kijas, H Kim, JK Lunney, FM McCarthy, JC McEwan, S Moore, B Nanduri, C Notredame, Y Palti, GS Plastow, JM Reecy, GA Rohrer, E Sarropoulou, CJ Schmidt, J Silverstein, RL Tellam, M Tixier-Boichard, G Tosser-Klopp, CK Tuggle, J Vilkki, SN White, S Zhao and H Zhou. Coordinated international action to accelerate genome-to-phenome with FAANG, the Functional Annotation of Animal Genomes project Genome Biology (2015) 16:57

Wang X, Lan X, Radunz AE, Khatib H. Maternal nutrition during pregnancy is associated with differential expression of imprinted genes and DNA methyltranfereases in muscle of beef cattle offspring. J Anim Sci. 2015 Jan;93(1):35-40.

Peñagaricano F, Wang X, Rosa GJ, Radunz AE, Khatib H. Maternal nutrition induces gene expression changes in fetal muscle and adipose tissues in sheep. BMC Genomics. 2014 Nov 28;15:1034.

Kropp J, Salih SM, Khatib H. 2014. Expression of microRNAs in bovine and human pre-implantation embryo culture media. Front Genet. 24;5:91. eCollection 2014.

Lan X, Cretney EC, Kropp J, Khateeb K, Berg MA, Peñagaricano F, Magness R, Radunz AE, Khatib H. 2013. Maternal Diet during Pregnancy Induces Gene Expression and DNA Methylation Changes in Fetal Tissues in Sheep. Front Genet 5;4:49. eCollection 2013.

Peñagaricano F, Weigel KA, Rosa GJ, Khatib H. 2013.Inferring quantitative trait pathways associated with bull fertility from a genome-wide association study. Front Genet. 3:307.

Khatib H. 2012. Bovine genomics by james e. Womack. Front Genet 3:275.

Peñagaricano F, Weigel KA, Khatib H. 2012. Genome-wide association study identifies candidate markers for bull fertility in Holstein dairy cattle. Anim Genet 43 Suppl 1:65-71.

Huang W, Long N, Khatib H. 2012. Genome-wide identification and initial characterization of bovine long non-coding RNAs from EST data. Anim Genet 43(6):674-82.Epub 2012 Feb 8.

Driver AM, Peñagaricano F, Huang W, Ahmad KR, Hackbart KS, Wiltbank MC, Khatib H. 2012. RNA-Seq analysis uncovers transcriptomic variations between morphologically similar in vivo- and in vitro-derived bovine blastocysts. BMC Genomics 13:118.

Peñagaricano F, Khatib H. 2012. Association of milk protein genes with fertilization rate and early embryonic development in Holstein dairy cattle. J Dairy Res 79(1):47-52. Epub 2011 Oct 13.

Huang W, Khatib H. 2010. Comparison of transcriptomic landscapes of bovine embryos using RNA-Seq. BMC Genomics 11:711.

Khatib H, Monson RL, Huang W, Khatib R, Schutzkus V, Khateeb H, Parrish JJ. 2010. Short communication: Validation of in vitro fertility genes in a Holstein bull population. J Dairy Sci 93(5):2244-9.

Huang W, Kirkpatrick BW, Rosa GJ, Khatib H. 2010. A genome-wide association study using selective DNA pooling identifies candidate markers for fertility in Holstein cattle. Anim Genet 41(6):570-8.

Zaitoun I, Downs KM, Rosa GJ, Khatib H. 2010. Upregulation of imprinted genes in mice: an insight into the intensity of gene expression and the evolution of genomic imprinting. Epigenetics 5(2):149-58. Epub 2010 Mar 1.

Huang W, Yandell BS, Khatib H. 2010. Transcriptomic profiling of bovine IVF embryos revealed candidate genes and pathways involved in early embryonic development. BMC Genomics 11:23.


Methods and compositions for improved cattle longevity and milk production
Publication date: 2014-06-05 Patent application number: US 14/040,757

Single nucleotide polymorphisms associated with bull fertility
Publication date: 2014-02-09 Patent application number: CA 2823638

Association of the progesterone receptor with fertility
Publication date: 2014-02-11 Patent application number: US 12/946,865

Genetic testing for improved cattle fertility
Publication date: 2013-10-10 Patent application number: US 13/798,176

Methods and compositions for improved cattle longevity and milk production
Publication date: 2013-06-06 Patent application number: PCT/US2008/082811

Methods and compositions for improved fertilization and embryonic survival
Publication date: 2012-05-31 Patent application number: US 13/276,333

Methods and compositions for genetically detecting improved milk production traits in cattle
Publication date: 2011-02-15 Patent application number: US 11/869,464

Methods and compositions for testing and breeding cattle for improved fertility and embryonic survival
Publication date: 2010-07-08 Patent application number: CA2743704

Detection of Lethality Gene for Improved Fertility in Mammals
Publication date: 2008-05-08 Patent application number: PCT/US2007/064103

Improved milk production in dairy cattle containing polymorphism in lectin-like oxidized ldl receptor (olri) gene
Publication date: 2007-01-13 Patent application number: CA2551173